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Instance-level image retrieval in fashion is a challenging issue owing to its increasing importance in real-scenario visual fashion search. Cross-domain fashion retrieval aims to match the unconstrained customer images as queries for photographs provided by retailers; however, it is a difficult task due to a wide range of consumer-to-shop (C2S) domain discrepancies and also considering that clothing image is vulnerable to various non-rigid deformations. To this end, we propose a novel multi-scale and multi-granularity feature learning network (MMFL-Net), which can jointly learn global-local aggregation feature representations of clothing images in a unified framework, aiming to train a cross-domain model for C2S fashion visual similarity. First, a new semantic-spatial feature fusion part is designed to bridge the semantic-spatial gap by applying top-down and bottom-up bidirectional multi-scale feature fusion. Next, a multi-branch deep network architecture is introduced to capture global salient, part-informed, and local detailed information, and extracting robust and discrimination feature embedding by integrating the similarity learning of coarse-to-fine embedding with the multiple granularities. Finally, the improved trihard loss, center loss, and multi-task classification loss are adopted for our MMFL-Net, which can jointly optimize intra-class and inter-class distance and thus explicitly improve intra-class compactness and inter-class discriminability between its visual representations for feature learning. Furthermore, our proposed model also combines the multi-task attribute recognition and classification module with multi-label semantic attributes and product ID labels. Experimental results demonstrate that our proposed MMFL-Net achieves significant improvement over the state-of-the-art methods on the two datasets, DeepFashion-C2S and Street2Shop.